Terrestrial adaptation refers to the various physiological and morphological changes that organisms undergo to thrive in land-based environments. These adaptations are crucial for overcoming challenges such as desiccation, gravity, and nutrient acquisition, allowing plants to survive and reproduce outside of aquatic habitats. In the context of bryophytes, which are non-vascular plants like mosses, liverworts, and hornworts, these adaptations are essential as they represent some of the first steps toward life on land.