P680 is a chlorophyll a molecule found in the reaction center of Photosystem II, which plays a crucial role in the light-dependent reactions of photosynthesis. This pigment absorbs light energy most efficiently at a wavelength of 680 nanometers, which is essential for the process of converting light energy into chemical energy. P680 acts as a primary electron donor, becoming oxidized when it absorbs light, leading to the production of ATP and NADPH.